Yup, and since the seat is conical, unless something miraculous happens, it will bear that load multi-axial. Meaning (drum roll please) that the lugs play at least a part in holding wheel centered to the hub.
Based on the equations I furnished, which the other recognized as worthwhile enough to site when they saw fit, it is clear that the studs and nuts can hold the wheel centered. Even the "Senior Axle Engineer" said that the studs don't flex, so what is the point of centering rings?